...
A contact is considered linked with an opportunity when they are in the Contact Roles section of the opportunity. Opportunity criteria applies to all Contacts in the Contact Roles section. For example, suppose you had an opportunity worth $100,000 with two people in the Contact Roles section, one with a job title of VP of Sales and one as VP of Marketing. An automation using opportunity criteria of Has Opportunity with > $99,000 in revenue would select both people. However, if the criteria Job Title contains "Sales" was added with the opportunity criteria, only the VP of Sales would be selected. In summary, Lead Liaison links contacts to opportunities through the Contact Role component of an opportunity:
...